Many of these primitive civilizations did not get beyond distinguishing among one, two, and many; others _______, and were able to operate with, large whole numbers. Still others achieved the recognition of numbers as abstract concepts, the adop­tion of special words for the individual numbers, the introduction of symbols for numbers, and even the use of a base such as ten, twenty, or five to denote a larger unit of quantity.

Which of the following BEST completes the blank?

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