There are birds that migrate by the stars, bats that echolocate, bees that compute the variance of flower patches, spiders that spin webs, humans that speak, ants that farm, lions that hunt in teams, cheetahs that hunt alone, monogamous gibbons, polyandrous seahorses, polygynous gorillas.

There are millions of animal species on earth, each with a different set of cognitive programs. The same basic neural tissue embodies all of these programs, and it could support many others as well. Facts about the properties of neurons, neurotransmitters, and cellular development cannot tell you which of these millions of programs the human mind contains. Even if all neural activity is the expression of a uniform process at the cellular level, it is the arrangement of neurons—into bird song templates or web-spinning programs—that matters.

The word “expression” in the passage is closest in meaning to?

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