Context of Chief

It isn’t known how these genes may relate to vegetarianism. But one of the chief differences between animal-based foods and plant-based ones is the chemical make-up of their fats or oils, collectively known as lipids. Yaseen and his colleagues speculate that some people may function better on a vegetarian diet because they are more able to synthesize certain lipid molecules that are present in meat.
–New Scientist

Which of the following BEST interprets the meaning of “chief” in the passage?

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