Through generations of close observation, they determined how to weed, prune, irrigate, transplant, and otherwise manipulate their environments to favor plants that provided food and medicine. They also developed specialized tools for digging and grinding as well as more effective methods of drying and storing seeds. The most sophisticated early plant cultivators lived in Mesoamerica, where maize agriculture was highly developed by 2500 b.c.e..

The word “sophisticated” in the passage is closest in meaning to? 


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